They are sentient, yet there are times when they are piloted by Shadow Angels. The Shadow Angels facilitate their harvesting via giant, floating harvesting machines called harvest beasts ( 収穫獣 shūkakujū ?) which are guarded by giant mecha called Cherubim Soldiers ( ケルビム兵 Kerubimu Hei ?). The Shadow Angels extract the prana ( プラーナ purāna ?, life force) from the harvested humans to serve as energy and nutrition to them, but also to feed the legendary Tree of Life ( 生命の樹 Inochi no Ki ?). The Shadow Angels begin invading human cites on Earth to harvest human beings, dubbing them as "the wingless ones", like cattle. They have been in slumber for twelve thousand years in their city of Atlandia ( アトランディア Atorandia ?) and have technology so advanced, it may be defined as very nearly magic.
Along with the disaster came the appearance of the Shadow Angels ( 堕天翅 Datenshi ?), winged beings with unexplainable powers and do not die of old age. Eleven years before the start of the series, a disaster only referred to as the "Great Catastrophe" destroyed/devastated significant parts of the world and killed off many of its inhabitants.
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